wo worlds exist, Sylvarant and Tethe'alla, each unaware of the existence of the other. In order for one world to flourish, the other must deteriorate. From each world, a Chosen is sent on a journey to restore that world's mana by awakening the spirits and becoming an angel for the sake of the goddess Martel.

Colette Brunel, the Chosen of Sylvarant, sets out on her journey accompanied by her best friends Lloyd Irving and Genis Sage. As they travel, they meet new friends and enemies, while they learn more about the truth behind the World Regeneration